the growing structure. The other two, the generative nuclei, can be
thought of as nonmotile sperm cells. After reaching an ovule and
breaking out of the pollen tube tip, one generative nucleus unites with
the egg cell to form a diploid zygote . The zygote undergoes a limited
number of divisions and gives rise to an embryo. The other generative
nucleus fuses with the two polar nuclei to produce a triploid nucleus,
which divides repeatedly before cell-wall formation occurs. This
process gives rise to the triploid endosperm, a nutrient tissue that
contains a variety of storage materials—such as stratch, sugars, fats,
proteins, hemicelluloses and and phytate (a phosphate reserve). The
events just described constitute what is called the double-
fertilization process, one of the characteristic features of all flowering
plants. In the orchids and in some other plants with minute seeds that
contain no reserve materials, endosperm formation is completely
suppressed. In other cases it is greatly reduced, but the reserve
materials are present elsewhere e.g., in the cotyledons, or seed leaves,
of the embryo, as in beans, lettuce, and peanuts, or in a tissue derived
from the nucellus, the perisperm, as in coffee. Other seeds, such as
those of beets, contain both perisperm and endosperm. The seed coat,
or testa, is derived from the one or two protective integuments of the
ovule. The ovary, in the simplest case, develops into a fruit. In many
plants, such as grasses and lettuce, the outer integument and ovary
wall are completely fused, so seed and fruit form one entity; such
seeds and fruits can logically be described together as “dispersal
units,” or diaspores. More often, however, the seeds are discrete units