STUDY OF HUMAN EVOLUTION NIRBHAY SUDHIR PIMPLE DEPARTMENT OF ZOOLOGY ABASAHEB GARWARE COLLEGE KARVE ROAD, PUNE-4.
Evolution is change in the heritable characteristics of biological populations over successive generations . These characteristics are the expressions of genes that are passed on from parent to offspring during reproduction . Different characteristics tend to exist within any given population as a result of mutation , genetic recombination and other sources of genetic variation . Evolution occurs when evolutionary processes such as natural selection (including sexual selection ) and genetic drift act on this variation, resulting in certain characteristics becoming more common or rare within a population. What is evolution?
The scientific theory of evolution by natural selection was conceived independently by Charles Darwin and Alfred Russel Wallace in the mid-19th century and was set out in detail in Darwin's book the Origin of Species . Human evolution is the evolutionary process that led to the emergence of anatomically modern humans , beginning with the evolutionary history of primates —in particular genus Homo —and leading to the emergence of Homo sapiens as a distinct species of the hominid family, which includes the great apes. This process involved the gradual development of traits such as human bipedalism and language . As well as interbreeding with other hominins , which indicate that human evolution was not linear but a web.
Australopithecus
Australopithecus , (Latin: “southern ape”) (genus Australopithecus ), group of extinct primates closely related to, if not actually ancestors of, modern human beings and known from a series of fossils found at numerous sites in eastern, north-central, and southern Africa . The various species of Australopithecus lived 4.4 million to 1.4 million years ago ( mya ), during the Pliocene and Pleistocene epochs (which lasted from 5.3 million to 11,700 years ago). DESCRIPTION
As characterized by the fossil evidence, members of Australopithecus bore a combination of human like and ape like traits. They were similar to modern humans in that they were bipedal (that is, they walked on two legs), but, like apes , they had small brains . Their canine teeth were smaller than those found in apes, and their cheek teeth were larger than those of modern humans. The general term australopith (or australopithecine ) is used informally to refer to members of the genus Australopithecus . Australopithecines include the genus Paranthropus (2.3–1.2 mya ), which comprises three species of australopiths —collectively called the “ robusts ” because of their very large cheek teeth set in massive jaws. CHARECTERS
Homo erectus
Homo erectus , (Latin: “upright man”) extinct species of the human genus ( Homo ), perhaps an ancestor of modern humans ( Homo sapiens ). H. erectus most likely originated in Africa , though Eurasia cannot be ruled out. Regardless of where it first evolved, the species seems to have dispersed quickly, starting about 1.9 million years ago ( mya ) near the middle of the Pleistocene Epoch , moving through the African tropics, Europe , South Asia , and Southeast Asia DESCRIPTION
The first fossils attributed to Homo erectus were discovered by a Dutch army surgeon, Eugène Dubois , who began his search for ancient human bones on the island of Java (now part of Indonesia) in 1890. Dubois found his first specimen in the same year, and in 1891 a well-preserved skullcap was unearthed at Trinil on the Solo River . Considering its prominent browridges , retreating forehead, and angled rear skull, Dubois concluded that the Trinil cranium showed anatomic features intermediate between those of humans (as they were then understood) and those of ap e s . DESCRIPTION
This history has been recorded directly if imprecisely by many sites that have yielded fossil remains of H. erectus . At other localities, broken animal bones and stone tools have indicated the presence of the species, though there are no traces of the people themselves. H. erectus was a human of medium stature that walked upright. The braincase was low, the forehead was receded, and the nose, jaws, and palate were wide. The brain was smaller and the teeth larger than in modern humans. H. erectus appears to have been the first human species to control fire , some 1,000,000 years ago. The species seems to have flourished until some 200,000 years ago (200 kya ) or perhaps later before giving way to other humans including Homo sapiens .
Several years later, near where the skull was discovered, he found a remarkably complete and modern-looking femur (thighbone). Since this bone was so similar to a modern human femur, Dubois decided that the individual to which it belonged must have walked erect. He adopted the name Pithecanthropus (coined earlier by the German zoologist Ernst Haeckel ) and called his discoveries Pithecanthropus erectus (“upright ape-man”), but the colloquial term became “ Java man .” col *lo*qui* al adjective \ kə -ˈ lō-kwē-əl \:used in or characteristic of familiar and informal conversation; also : unacceptably informal: using conversational style
Neanderthal man
Neanderthal , ( Homo neanderthalensis , Homo sapiens neanderthalensis ), also spelled Neandertal , member of a group of archaic humans who emerged at least 200,000 years ago during the Pleistocene Epoch (about 2.6 million to 11,700 years ago) and were replaced or assimilated by early modern human populations ( Homo sapiens ) between 35,000 and perhaps 24,000 years ago. Neanderthals inhabited Eurasia from the Atlantic regions of Europe eastward to Central Asia , from as far north as present-day DESCRIPTION
Belgium and as far south as the Mediterranean and southwest Asia. Similar archaic human populations lived at the same time in eastern Asia and in Africa. Because Neanderthals lived in a land of abundant limestone caves , which preserved bones well, and where there has been a long history of prehistoric research, they are better known than any other archaic human group. Consequently, they have become the archetypal “cavemen.” The name Neanderthal (or Neandertal ) derives from the Neander Valley (German Neander Thal or Neander Tal ) in Germany, where the fossils were first found.
Although Neanderthals possessed much in common physically with early modern humans, the constellation of Neanderthal features is unique, with much variation among individuals as far as craniofacial (head and facial) characteristics are concerned. Features of the cranium and lower jaw that were present more often in Neanderthals than in early and recent modern humans include a low-vaulted cranium, large orbital and nasal openings, and prominent arched brow ridges. A pronounced occipital region (the rear and base of the skull) served to anchor the large neck musculature.
The cranial capacity of Neanderthals was similar to or larger than that of recent humans. The front teeth were larger than those in modern humans, but the molars and premolars were of a similar size. The lower jaw displayed a receding chin and was robustly built. The mental foramen, a small hole in the skull that allows nerves to reach the lower jaw, was placed farther back in Neanderthals than in recent humans, and a space between the last molar and the ascending edge of the lower jaw occurred in many individuals. There was also apparently less lumbar lordosis (back curvature) in Neanderthals
HOMO SAPIENS SAPIENS
Homo sapiens , (Latin: “wise man”) the species to which all modern human beings belong. Homo sapiens is one of several species grouped into the genus Homo , but it is the only one that is not extinct . The name Homo sapiens was applied in 1758 by the father of modern biological classification Carolus Linnaeus . Traditionally, this subspecies designation was used by paleontologists and anthropologists to separate modern human beings from more-archaic members of Homo sapiens . H. s. sapiens is thought to have evolved sometime between 160,000 and 90,000 years ago in Africa before migrating first to the Middle East and Europe and later to Asia , Australia , and the Americas . DESCRIPTION
Currently, H. s. sapiens is the only widely accepted subspecies of H. sapiens , and the necessity of this designation remains a matter of debate, since traditional taxonomic practice subdivides a species only when there is evidence of two or more distinct subgroups. Several subspecies of H. sapiens have been proposed. For example, Swedish naturalist and explorer Carl Linnaeus , in the 10th edition of his Systema Naturae (1758), classified modern human beings into four subspecies according to geographic origin: H. s. asiaticus , H. s. europaeus , H. s. afer , and H. s. americanus . Linnaeus’s classification was later discarded because of the recognition of racial prejudice and outdated notions of European superiority implicit in his taxonomy and because of discoveries that only superficial differences existed between these groups
the earliest Homo sapiens had bodies with short, slender trunks and long limbs. These body proportions are an adaptation for surviving in tropical regions due to the greater proportion of skin surface available for cooling the body. More stocky builds gradually evolved when populations spread to cooler regions, as an adaptation that helped the body retain heat. Modern humans now have an average height of about 160 centimetres in females and 175 centimetres in males. Brain Homo sapiens living today have an average brain size of about 1350 cubic centimetres which makes-up 2.2% of our body weight. Early Homo sapiens , however, had slightly larger brains at nearly 1500 cubic centimetres .
Skull modern Homo sapiens skulls have a short base and a high braincase. Unlike other species of Homo , the skull is broadest at the top. The fuller braincase also results in almost no post-orbital constriction or narrowing behind the eye sockets back of the skull is rounded and indicates a reduction in neck muscles face is reasonably small with a projecting nose bone brow ridge is limited and the forehead is tall orbits (eye sockets) are square rather than round Jaws and teeth jaws are short which result in an almost vertical face usually no gap ( retromolar space) between the last molar teeth and the jaw bone jaws are lightly built and have a protruding bony chin for added strength. Homo sapiens is the only species to have a protruding chin. shortened jaw has affected the arrangement of the teeth within the jaw. They are now arranged in a parabolic shape in which the side rows of teeth splay outwards rather than remain parallel as in our earliest long jawed ancestors. teeth are relatively small compared with earlier species. This is especially noticeable in the front incisor and canine teeth. front premolar teeth in the lower jaw have two equal-sized cusps (bumps on the chewing surface